I didn't have any trouble importing it, using nothing but the "automatic bone orientation" setting. Even the animation came through. But I'mdon't know why exactly this is happening (I'm using 2.82a and not 2having the same issue), but since you have so few bones which are misaligned, I find that I was able to go into Edit Mode and manually adjust them in less than five minutes.81
Regretably, this is not the answer you were hoping for. Consider upgradingBut it seems to a later version of Blender and see if that makes a differencework.