Keyframed UVs
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As most of all the properties, UV island coordinates can be keyframed.

By examining the Outliner or the Graph editor you can see how each island has fixed location vector, so if you modify the UV island position and save without overriding the keframes values, the island would come back to the stored position.

[![enter image description here][1]][1]


  [1]: https://i.sstatic.net/QOovK.png