Keyframed UVs
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As most of all the properties, UV island coordinates can be individually keyframed.

By examining the **Outliner** and the **Graph editor** you can realize that each island has fixed location coordinates for the current animation.

[![enter image description here][1]][1]

If you modify the UV island position and save without overriding the keframes values, the island would come back to the stored position.

If those keframes are needed, you must **Insert** a new keframe upon the old one before saving.


  [1]: https://i.sstatic.net/QOovK.png